The App modules would have to be recompiled for the older gen 1 v4.x firmware because the older gen My Cloud uses a different page file size (64K if I remember right). Further there is no “define.js” in the v4.x firmware (that I can find by searching). And the HTML code of the rendered Dashboard under v4.x also does not have an “Apps” entry in the nav bar area of the code.
The better way for the older first gen My Cloud’s is do something like the following:
http://community.wd.com/t/clean-os-debian-openmediavault-and-other-firmwares/93714
Or if one wants Transmission alone:
http://community.wd.com/t/app-transmission-various-versions-for-firmware-v4-03-2016/93424
Or see the following:
http://community.wd.com/t/repository-with-software-worked-on-v4-firmware/94532
There are a number of prior threads for running Transmission under v3.x/v4.x firmware first gen My Clouds. One can use the forum search feature, magnifying glass icon upper right, to find those other threads.